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Beutel\uff06Richard A. Leschen\uff08\u7de8\u96c6\u8005\uff09\u30cf\u30f3\u30c9\u30d6\u30c3\u30af\u306e\u52d5\u7269\u5b66\u3002\u30dc\u30ea\u30e5\u30fc\u30e0IV\u3002\u7bc0\u8db3\u52d5\u7269\uff1a\u6606\u866b\u3002\u30d1\u30fc\u30c838.\u7532\u866b\u76ee\u3002\u7b2c1\u5dfb\uff1a\u5f62\u614b\u3068\u7cfb\u7d71\u5b66\u3001Archostemata\u3001Adephaga\u3001Myxophaga\u3001Polyphaga Partim\u3002\u30cb\u30e5\u30fc\u30e8\u30fc\u30af\u5dde\u30d9\u30eb\u30ea\u30f3\uff1a\u30a6\u30a9\u30eb\u30bf\u30fc\u30fb\u30c7\u30fb\u30b0\u30eb\u30fc\u30bf\u30fc\u3002 P. 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