[{"@context":"http:\/\/schema.org\/","@type":"BlogPosting","@id":"https:\/\/wiki.edu.vn\/en\/wiki40\/list-of-ground-beetle-genera\/#BlogPosting","mainEntityOfPage":"https:\/\/wiki.edu.vn\/en\/wiki40\/list-of-ground-beetle-genera\/","headline":"List of ground beetle genera","name":"List of ground beetle genera","description":"From Wikipedia, the free encyclopedia This page lists the genera in the family Carabidae, according to Catalogue of Life and","datePublished":"2021-06-26","dateModified":"2021-06-26","author":{"@type":"Person","@id":"https:\/\/wiki.edu.vn\/en\/wiki40\/author\/lordneo\/#Person","name":"lordneo","url":"https:\/\/wiki.edu.vn\/en\/wiki40\/author\/lordneo\/","image":{"@type":"ImageObject","@id":"https:\/\/secure.gravatar.com\/avatar\/c9645c498c9701c88b89b8537773dd7c?s=96&d=mm&r=g","url":"https:\/\/secure.gravatar.com\/avatar\/c9645c498c9701c88b89b8537773dd7c?s=96&d=mm&r=g","height":96,"width":96}},"publisher":{"@type":"Organization","name":"Enzyklop\u00e4die","logo":{"@type":"ImageObject","@id":"https:\/\/wiki.edu.vn\/wiki4\/wp-content\/uploads\/2023\/08\/download.jpg","url":"https:\/\/wiki.edu.vn\/wiki4\/wp-content\/uploads\/2023\/08\/download.jpg","width":600,"height":60}},"image":{"@type":"ImageObject","@id":"https:\/\/en.wikipedia.org\/wiki\/Special:CentralAutoLogin\/start?type=1x1","url":"https:\/\/en.wikipedia.org\/wiki\/Special:CentralAutoLogin\/start?type=1x1","height":"1","width":"1"},"url":"https:\/\/wiki.edu.vn\/en\/wiki40\/list-of-ground-beetle-genera\/","about":["Wiki"],"wordCount":20733,"articleBody":"From Wikipedia, the free encyclopediaThis page lists the genera in the family Carabidae, according to Catalogue of Life and Carabcat Database as of March 2023, excluding Tiger Beetles (which are considered to be in the family Cicindelidae).[1][2]Table of ContentsFamily Carabidae[edit]Subfamily Anthiinae Bonelli, 1813[edit]Subfamily Apotominae LeConte, 1853[edit]Subfamily Brachininae Bonelli, 1810[edit]Subfamily Broscinae Hope, 1838[edit]Subfamily Carabinae Linnaeus, 1802[edit]Subfamily Ctenodactylinae Laporte, 1834[edit]Subfamily Dryptinae Bonelli, 1810[edit]Subfamily Elaphrinae Latreille, 1802[edit]Subfamily Gineminae Ball & Shpeley, 2002[edit]Subfamily Harpalinae Bonelli, 1810[edit]Subfamily Hiletinae Schi\u00f6dte, 1847[edit]Subfamily Lebiinae Bonelli, 1810[edit]Subfamily Licininae Bonelli, 1810[edit]Subfamily Loricerinae Bonelli, 1810[edit]Subfamily Melaeninae Csiki, 1933[edit]Subfamily Migadopinae Chaudoir, 1861[edit]Subfamily Nebriinae Laporte, 1834[edit]Subfamily Nototylinae B\u00e4nninger, 1927[edit]Subfamily Omophroninae Bonelli, 1810[edit]Subfamily Orthogoniinae Schaum, 1857[edit]Subfamily Panagaeinae Bonelli, 1810[edit]Subfamily Patrobinae Kirby, 1837[edit]Subfamily Paussinae Latreille, 1806[edit]Subfamily Platyninae Bonelli, 1810[edit]Subfamily Promecognathinae LeConte, 1853[edit]Subfamily Pseudomorphinae Hope, 1838[edit]Subfamily Psydrinae LeConte, 1853[edit]Subfamily Pterostichinae Bonelli, 1810[edit]Subfamily Rhysodinae Laporte, 1840[edit]Subfamily Scaritinae Bonelli, 1810[edit]Subfamily Siagoninae Bonelli, 1813[edit]Subfamily Trechinae Bonelli, 1810[edit]Subfamily Xenaroswellianinae Erwin, 2007[edit]Subfamily \u2020Conjunctiinae Ponomarenko, 1977[edit]Subfamily \u2020Protorabinae Ponomarenko, 1977[edit]References[edit]Family Carabidae[edit]Subfamily Anthiinae Bonelli, 1813[edit]Tribe Anthiini Bonelli, 1813Anthia Weber, 1801Atractonotus Perroud, 1847Baeoglossa Chaudoir, 1850Cycloloba Chaudoir, 1850Cypholoba Chaudoir, 1850Eccoptoptera Chaudoir, 1878Gonogenia Chaudoir, 1844Netrodera Chaudoir, 1850Tribe Helluonini Hope, 1838Subtribe Helluonina Hope, 1838Aenigma Newman, 1836Ametroglossus Sloane, 1914Dicranoglossus Chaudoir, 1872Epimicodema Sloane, 1914Gigadema J.Thomson, 1859Helluapterus Sloane, 1914Helluarchus Sloane, 1914Helluo Bonelli, 1813Helluodema Laporte, 1867Helluonidius Chaudoir, 1872Helluopapua Darlington, 1968Helluosoma Laporte, 1867Neohelluo Sloane, 1914Platyhelluo Baehr, 2005Subtribe Omphrina Jedlicka, 1941Colfax Andrewes, 1920Creagris Nietner, 1857Dailodontus Reiche, 1843Erephognathus Alluaud, 1932Helluobrochus Reichardt, 1974Helluomorpha Laporte, 1834Helluomorphoides Ball, 1951Macrocheilus Hope, 1838Meladroma Motschulsky, 1855Omphra Dejean, 1825Pleuracanthus Gray, 1832Triaenogenius Chaudoir, 1877Tribe Physocrotaphini Chaudoir, 1863Anguloderus Anichtchenko & Sciaky, 2017Foveocrotaphus Anichtchenko, 2014Helluodes Westwood, 1847Holoponerus Fairmaire, 1883Physocrotaphus Parry, 1849Physoglossus Akhil & Sabu, 2020Pogonoglossus Chaudoir, 1863Schuelea Baehr, 2004Subfamily Apotominae LeConte, 1853[edit]Apotomus Illiger, 1807Subfamily Brachininae Bonelli, 1810[edit]Tribe Brachinini Bonelli, 1810Aptinoderus Hubenthal, 1919Aptinus Bonelli, 1810Brachinulus Basilewsky, 1958Brachinus Weber, 1801Mastax Fischer von Waldheim, 1828Pheropsophus Solier, 1833Styphlodromus Basilewsky, 1959Styphlomerus Chaudoir in Putzeys, 1875Tribe Crepidogastrini Jeannel, 1949Brachynillus Reitter, 1904Crepidogaster Boheman, 1848Crepidogastrillus Basilewsky, 1959Crepidogastrinus Basilewsky, 1957Crepidolomus Basilewsky, 1959Crepidonellus Basilewsky, 1959Subfamily Broscinae Hope, 1838[edit]Tribe Broscini Hope, 1838Subtribe Axonyina Roig-Ju\u00f1ent, 2000Broscodes Bolivar y Pieltain, 1914Rawlinsius Davidson & Ball, 1998Subtribe Baripodina Jeannel, 1941Baripus Dejean, 1828Bembidiomorphum Champion, 1918Subtribe Broscina Hope, 1838Broscodera Lindroth, 1961Broscosoma Rosenhauer, 1846Broscus Panzer, 1813Chaetobroscus Semenov, 1900Craspedonotus Schaum, 1863Eobroscus Kryzhanovskij, 1951Kashmirobroscus J.Schmidt; Wrase & Sciaky, 2013Miscodera Eschscholtz, 1830Zacotus LeConte, 1869Subtribe Creobiina Jeannel, 1941Acallistus Sharp, 1886Adotela Laporte, 1867Anheterus Putzeys, 1868Bountya Townsend, 1971Brithysternum W.J.MacLeay, 1873Cascellius Curtis, 1838Cerotalis Laporte, 1867Creobius Gu\u00e9rin-M\u00e9neville, 1838Gnathoxys Westwood, 1842Nothocascellius Roig-Ju\u00f1ent, 1995Promecoderus Dejean, 1829Subtribe Nothobroscina Roig-Ju\u00f1ent, 2000Chylnus Sloane, 1920Diglymma Sharp, 1886Eurylychnus Bates, 1891Mecodema Blanchard, 1843Monteremita Seldon & Holwell, 2019Nothobroscus Roig-Ju\u00f1ent & Ball, 1995Oregus Putzeys, 1868Orthoglymma Liebherr; Marris; Emberson; Syrett & Roig-Ju\u00f1ent, 2011Percolestus Sloane, 1892Percosoma Schaum, 1858Subfamily Carabinae Linnaeus, 1802[edit]Tribe Carabini Linnaeus, 1802Subtribe Carabina Linnaeus, 1802Aplothorax G.R.Waterhouse, 1842Calosoma Weber, 1801Carabus Linnaeus, 1758Subtribe Ceroglossina Lapouge, 1927Ceroglossus Solier, 1848Tribe Cychrini Perty, 1830Subtribe Cychrina Perty, 1830Cychropsis Boileau, 1901Cychrus Fabricius, 1794Scaphinotus Dejean, 1826Sphaeroderus Dejean, 1826Subtribe Pamborina Hope, 1838Maoripamborus Brookes, 1944Pamborus Latreille, 1812Subfamily Ctenodactylinae Laporte, 1834[edit]Tribe Ctenodactylini Laporte, 1834Alachnothorax Liebke, 1929Amblycoleus Chaudoir, 1872Antipionycha Liebke, 1928Askalaphium Liebke, 1938Ctenodactyla Dejean, 1825Leptotrachelon Liebke, 1928Leptotrachelus Latreille, 1829Oilea Liebke, 1931Parapionycha Liebke, 1929Pionycha Chaudoir, 1848Plagiotelum Solier, 1849Propionycha Liebke, 1928Pseudometabletus Liebke, 1930Schidonychus Klug, 1834Teukrus Liebke, 1931Wate Liebke, 1928Tribe Hexagoniini G.Horn, 1881Dinopelma Bates, 1889Hexagonia Kirby, 1825Omphreoides Fairmaire, 1896Subfamily Dryptinae Bonelli, 1810[edit]Tribe Dryptini Bonelli, 1810Desera Hope, 1831Drypta Latreille, 1797Megadrypta Sciaky & Anichtchenko, 2020Neodrypta Basilewsky, 1960Nesiodrypta Jeannel, 1949Prionodrypta Jeannel, 1949Tribe Galeritini Kirby, 1825Ancystroglossus Chaudoir, 1863Eunostus Laporte, 1835Galerita Fabricius, 1801Trichognatha Latreille, 1829Tribe Zuphiini Bonelli, 1810Subtribe Dicrodontina Machado, 1992Dicrodontus Chaudoir, 1872Subtribe Leleupidiina Basilewsky, 1951Colasidia Basilewsky, 1954Gunvorita Landin, 1955Leleupidia Basilewsky, 1951Neoleleupidia Basilewsky, 1953Paraleleupidia Basilewsky, 1951Subtribe Planetina Jedlicka, 1941Planetes W.S.MacLeay, 1825Subtribe Zuphiina Bonelli, 1810Acrogenys W.J.MacLeay, 1864Agastus Schmidt-Goebel, 1846Chaudoirella Mateu, 1982Coarazuphium Gnaspini; Vanin & Godoy, 1998Ildobates Espa\u00f1ol, 1966Metaxidius Chaudoir, 1852Metazuphium Mateu, 1992Mischocephalus Chaudoir, 1863Parazuphium Jeannel, 1942Polistichus Bonelli, 1810Pseudaptinus Laporte, 1834Speothalpius B.Moore, 1995Speozuphium B.Moore, 1995Typhlozuphium Baehr, 2014Zuphioides Ball & Shpeley, 2013Zuphium Latreille, 1805Subfamily Elaphrinae Latreille, 1802[edit]Blethisa Bonelli, 1810Diacheila Motschulsky, 1844Elaphrus Fabricius, 1775\u2020Elaphrotites Haupt, 1856Subfamily Gineminae Ball & Shpeley, 2002[edit]Ginema Ball & Shpeley, 2002Subfamily Harpalinae Bonelli, 1810[edit]Tribe Anisodactylini Lacordaire, 1854Allendia Noonan, 1974Allocinopus Broun, 1903Amphasia Newman, 1838Anisodactylus Dejean, 1829Anisostichus Emden, 1953Cenogmus Sloane, 1898Chydaeus Chaudoir, 1854Crasodactylus Gu\u00e9rin-M\u00e9neville, 1847Criniventer Emden, 1953Diachromus Erichson, 1837Dicheirus Mannerheim, 1843Gaioxenus Broun, 1910Geopinus LeConte, 1847Gnathaphanus W.S.MacLeay, 1825Gynandromorphus Dejean, 1829Haplaner Chaudoir, 1878Harpalomimetes Schauberger in Csiki, 1932Hypharpax W.S.MacLeay, 1825Maoriharpalus Larochelle & Larivi\u00e8re, 2005Nornalupia Kataev, 2002Notiobia Perty, 1830Parabaris Broun, 1881Progonochaetus G.M\u00fcller, 1938Pseudanisotarsus Noonan, 1973Pseudognathaphanus Schauberger, 1932Pseudorhysopus Kataev & Wrase, 2001Rhysopus Andrewes, 1929Scybalicus Schaum, 1862Triplosarus Bates, 1874Tuiharpalus Larochelle & Larivi\u00e8re, 2005Xestonotus LeConte, 1853Tribe Harpalini Bonelli, 1810Subtribe Amblystomina Fauvel, 1889Amblystomus Erichson, 1837Anomostomus LaFert\u00e9-S\u00e9nect\u00e8re, 1853Barysomus Dejean, 1829Oosoma Nietner, 1857Subtribe Ditomina Bonelli, 1810Bronislavia Semenov, 1891Carenochyrus Solsky, 1874Carterus Dejean, 1830Chilotomus Chaudoir, 1842Ditomus Bonelli, 1810Dixus Billberg, 1820Eocarterus Stichel, 1923Eucarterus Reitter, 1900Graniger Motschulsky, 1864Indocarterus Kataev & Wrase, 2018Liochirus Tschitscherine, 1897Machozetus Chaudoir, 1850Odotoncarus Solier, 1835Oedesis Motschulsky, 1850Pachycarus Solier, 1835Parapenthes Kataev & Wrase, 2018Penthus Chaudoir, 1843Phorticosomus Schaum, 1863Proditomus Schauberger, 1934Pseudaristus Reitter, 1900Tschitscherinellus Csiki, 1906Subtribe Harpalina Bonelli, 1810Acinopus Dejean, 1821Afromizonus Basilewsky, 1947Allosiopelus N.Ito, 1995Amblygnathus Dejean, 1829Anisocnemus Chaudoir, 1843Athrostictus Bates, 1878Axinotoma Dejean, 1829Aztecarpalus Ball, 1970Bleusea Bedel, 1897Boeomimetes P\u00e9ringuey, 1896Bradybaenus Dejean, 1829Coleolissus Bates, 1892Cratacanthus Dejean, 1829Cratognathus Dejean, 1829Daptus Fischer von Waldheim, 1823Dioryche W.S.MacLeay, 1825Discoderus LeConte, 1853Dregus Motschulsky, 1864Ectinothorax Alluaud, 1941Eriophonus Tschitscherine, 1901Euryderus LeConte, 1846Geodromus Dejean, 1829Harpalinus Jeannel, 1946Harpaliscus Bates, 1892Harpalobrachys Tschitscherine, 1899Harpalodiodes Bousquet, 2002Harpalomimus Facchini & Giachino, 2020Harpalomorphus P\u00e9ringuey, 1896Harpalus Latreille, 1802Harpathaumas Basilewsky, 1947Harponixus Basilewsky, 1950Hartonymus Casey, 1914Heteracantha Brull\u00e9, 1835Hyphaereon W.S.MacLeay, 1825Indiophonus N.Ito, 1996Kareya Andrewes, 1919Lampetes Andrewes, 1940Laparhetes Jeannel, 1946Liodaptus Bates, 1889Meroctenus Gemminger & Harold, 1868Microderes Faldermann, 1836Neoaulacoryssus Noonan, 1985Neodiachipteryx Noonan, 1985Neohyparpalus Clarke, 1981Neophygas Noonan, 1976Nesacinopus Tschitscherine, 1900Nesarpalus Bedel, 1897Nipponoharpalus Habu, 1973Nothodaptus Maindron, 1906Oesyperus Andrewes, 1923Omostropus P\u00e9ringuey, 1896Ooidius Chaudoir, 1847Ophoniscus Bates, 1892Ophonus Dejean, 1821Oxycentrus Chaudoir, 1854Panagrius Andrewes, 1933Pangus Dejean, 1821Parasiopelus Basilewsky, 1946Paraulacoryssus Shpeley; Hunting & Ball, 2017Parophonus Ganglbauer, 1891Paulianoscirtus Basilewsky, 1976Penthophonus Reitter, 1900Piosoma LeConte, 1847Platymetopsis Ball & Maddison, 1987Platymetopus Dejean, 1829Prakasha Andrewes, 1919Pseudodiachipteryx Burgeon, 1936Pseudohyparpalus Basilewsky, 1946Pseudoselenophorus P\u00e9ringuey, 1896Selenophorus Dejean, 1829Selenotichnus Kataev, 1999Siopelophonus Kataev, 2015Siopelus Murray, 1859Stenomorphus Dejean, 1831Trichopselaphus Chaudoir, 1843Trichotichnus A.Morawitz, 1863Trichoxycentrus N.Ito, 2000Typsiharpalus Tschitscherine, 1901Tribe Pelmatellini Bates, 1882Hakaharpalus Larochelle & Larivi\u00e8re, 2005Kupeharpalus Larochelle & Larivi\u00e8re, 2005Lecanomerus Chaudoir, 1850Nemaglossa Solier, 1849Notospeophonus B.Moore, 1962Pelmatellus Bates, 1882Syllectus Bates, 1878Trachysarus Reed, 1874Tribe Stenolophini Kirby, 1837Acupalpus Latreille, 1829Agonoleptus Casey, 1914Amerinus Casey, 1884Angionychus Klug, 1853Anthracus Motschulsky, 1850Batoscelis Dejean, 1836Bradycellus Erichson, 1837Cratosoma Jeannel, 1948Cyptomicrus Vinson, 1939Dicheirotrichus Jacquelin du Val, 1855Egadyla Alluaud, 1916Euthenarus Bates, 1874Fuminoria Morita, 2006Goniocellus Casey, 1914Gugheorites Basilewsky, 1951Haplanister B.Moore, 1996Hemiaulax Bates, 1892Hippoloetis Laporte, 1835Idiomelas Tschitscherine, 1900Kaffovatus Clarke, 1972Kenyacus Alluaud, 1917Kiwiharpalus Larochelle & Larivi\u00e8re, 2005Lioholus Tschitscherine, 1897Loxoncus Schmidt-Goebel, 1846Pachytrachelus Chaudoir, 1852Parabradycellus N.Ito, 2003Paramecus Dejean, 1829Philodes LeConte, 1861Pholeodytes Britton, 1962Pogonodaptus G.Horn, 1881Polpochila Solier, 1849Psychristus Andrewes, 1930Rhabidius Basilewsky, 1948Stenolophus Dejean, 1821Uenanthracus Kasahara, 1994Subfamily Hiletinae Schi\u00f6dte, 1847[edit]Eucamaragnathus Jeannel, 1938Hiletus Schi\u00f6dte, 1847Subfamily Lebiinae Bonelli, 1810[edit]Tribe Cyclosomini Laporte, 1834Subtribe Corsyrina GanglbauerCorsyra Dejean, 1825Discoptera Semenov, 1889Subtribe Cyclosomina Laporte, 1834Cyclosomus Latreille, 1829Sarothrocrepis Chaudoir, 1850Tetragonoderus Dejean, 1829Subtribe Graphipterina Latreille, 1802Graphipterus Latreille, 1802Piezia Brull\u00e9, 1834Trichopiezia N\u00e8gre, 1955Subtribe Masoreina Chaudoir, 1871Anaulacus W.S.MacLeay, 1825Atlantomasoreus Mateu, 1984Leuropus Andrewes, 1947Lophidius Dejean, 1831Masoreus Dejean, 1821Mnuphorus Chaudoir, 1873Odontomasoreus Darlington, 1968Somoplatodes Basilewsky, 1986Somoplatus Dejean, 1829Subtribe Nemotarsina Bates, 1883Nemotarsus LeConte, 1853Tribe Lachnophorini LeConte, 1853Subtribe Calophaenina Jeannel, 1948Calophaena Klug, 1821Calophaenoidea Liebke, 1930Subtribe Lachnophorina LeConte, 1853Amphithasus Bates, 1871Anchonoderus Reiche, 1843Aporesthus Bates, 1871Asklepia Liebke, 1938Balligratus Moret & Ortu\u00f1o, 2017Calybe Laporte, 1834Ega Laporte, 1834Eucaerus LeConte, 1853Euphorticus G.Horn, 1881Grundmannius Basilewsky, 1965Guatemalteca Erwin, 2004Lachnaces Bates, 1872Lachnophorus Dejean, 1831Peruphorticus Erwin & Zamorano, 2014Pseudophorticus Erwin, 2004Selina Motschulsky, 1858Tribe Lebiini Bonelli, 1810Subtribe Agrina Kirby, 1837Abaditicus Ball & Hilchie, 1983Abrodiella Bousquet, 2002Agra Fabricius, 1801Allardina Basilewsky, 1963Amelus Chaudoir, 1873Anomotariella Baehr, 2012Anomotarus Chaudoir, 1875Apterodromites Mateu, 1976Aspasiola Chaudoir, 1877Calleida Latreille, 1824Calleidomorpha Motschulsky, 1855Callidadelpha Steinheil, 1875Chaudoirina Mateu, 1955Cryptobatis Eschscholtz, 1829Cyanotarus Reed, 1874Cylindronotum Putzeys, 1845Demetrida White, 1846Diabaticus Bates, 1878Do Baehr, 2009Dromidea Perroud & Montrouzier, 1864Epikastea Liebke, 1935Eujalmenus Bousquet, 2002Euplatia Chaudoir, 1873Eurycalleida Maindron, 1905Falsodromius Mateu, 1976Glycia Chaudoir, 1842Hyboptera Chaudoir, 1873Hybopteroides Erwin & Ball, 2012Infernophilus Larson, 1969Kteatus Liebke, 1936Lipostratia Chaudoir, 1873Merizomena Chaudoir, 1873Mimodromius Chaudoir, 1873Mimophilorhizus Mateu, 1993Oechalius Liebke, 1935Ogygium Liebke, 1935Onota Chaudoir, 1873Otoglossa Chaudoir, 1873Paraglycia Bedel, 1904Phacocerus Chaudoir, 1873Philophuga Motschulsky, 1859Phloeocarabus W.J.MacLeay, 1871Plochionus Dejean, 1821Pontonoa Liebke, 1935Pseudotoglossa Mateu, 1961Pylartesius Liebke, 1939Rugitarus Baehr, 2009Speotarus B.Moore, 1964Straneotia Mateu, 1961Tecnophilus Chaudoir, 1877Teiresia Liebke, 1935Thoasia Liebke, 1939Titaresius Liebke, 1935Trigonothops W.J.MacLeay, 1864Valeriaaschero Erwin, 2004Vianasia Mateu, 1955Subtribe Apenina BallApenes LeConte, 1851Cymindoidea Laporte, 1833Habutarus Ball & Hilchie, 1983Platytarus Fairmaire, 1850Trymosternus Chaudoir, 1873Subtribe Celaenephina Habu, 1982Celaenephes Schmidt-Goebel, 1846Subtribe Cymindidina Laporte, 1834Afrotarus Jeannel, 1949Assadecma Basilewsky, 1982Cymindis Latreille, 1805Hystrichopus Boheman, 1848Leptosarcus P\u00e9ringuey, 1896Metaxymorphus Chaudoir, 1850Petrimagnia Kryzhanovskij & Mikhailov, 1971Pinacodera Schaum, 1857Plagiopyga Boheman, 1848Pseudomasoreus Desbrochers des Loges, 1904Taridius Chaudoir, 1875Subtribe Demetriadina Bates, 1886Cylindrocranius Chaudoir, 1878Demetrias Bonelli, 1810Subtribe Dromiusina Bonelli, 1810Afrodromius Basilewsky, 1958Axinopalpus LeConte, 1846Barrymooreana Baehr, 1997Brachynopterus Bedel, 1898Brigalowia Baehr, 2006Calodromius Reitter, 1905Carbonellia Mateu, 1968Disciferella Kataev & Muilwijk, 2020Dromiops P\u00e9ringuey, 1898Dromius Bonelli, 1810Dromoceryx Schmidt-Goebel, 1846Geoffreyella Baehr, 2012Klepteromimus P\u00e9ringuey, 1898Mesolestes Schatzmayr, 1943Metadromius Bedel, 1907Microdaccus Schaum in Baudi di Selve, 1864Microlestes Schmidt-Goebel, 1846Microlestodes Baehr, 1987Monnea Mateu, 1970Negrea Mateu, 1968Omophagus Andrewes, 1937Oxoides Solier, 1849Paradromius Fowler, 1887Philorhizus Hope, 1838Polyaulacus Chaudoir, 1878Psammodromius Peyerimhoff, 1927Pseudomonnea Mateu, 1983Somalodromius Mateu, 1967Xenodromius Bates, 1891Zolotarevskyella Mateu, 1953Subtribe Gallerucidiina Chaudoir, 1872Gallerucidia Chaudoir, 1872Lebidia A.Morawitz, 1862Subtribe Lebiina Bonelli, 1810Alkestis Liebke, 1939Aristolebia Bates, 1892Daer Semenov & Znojko, 1929Lachnolebia Maindron, 1905Lebia Latreille, 1802Lebidema Motschulsky, 1864Lebiomorphica Lorenz, 1998Lebistina Motschulsky, 1864Lebistinida P\u00e9ringuey, 1898Lionedya Chaudoir, 1871Matabele P\u00e9ringuey, 1896Megalebia Mateu, 1972Orthobasis Chaudoir, 1871Pachylebia Jeannel, 1949Pachylebiodes Mateu, 1972Paulianolebia Mateu, 1972Pseudopachylebia Mateu, 1972Rhopalostyla Chaudoir, 1850Scalidion Schmidt-Goebel, 1846Setolebia Jedlicka, 1941Sofota Jedlicka, 1951Subtribe Lichnasthenina J.ThomsonAstastus P\u00e9ringuey, 1896Australovelinda Baehr, 2012Imasakaia Arai & Morita, 2010Lichnasthenus J.Thomson, 1858Mimovelindopsis Mateu, 1963Velinda Andrewes, 1921Velindastus Sch\u00fcle & Lorenz, 2008Velindomimus Jeannel, 1955Velindopsis Burgeon, 1937Subtribe Lionychina Jeannel, 1948Apristomimus Mateu, 1969Apristus Chaudoir, 1846Eremolestes Maindron, 1905Lionychus Wissmann, 1846Lorestania Anichtchenko, 2011Metablus Jedlicka, 1958Omobrus Andrewes, 1930Singiliomimus P\u00e9ringuey, 1896Syntomus Hope, 1838Tilius Chaudoir, 1876Trichidema Basilewsky, 1956Subtribe Metallicina Basilewsky, 1984Euproctinus Leng & Mutchler, 1927Metallica Chaudoir, 1873Pachycallida Jeannel, 1949Parena Motschulsky, 1860Subtribe Peliocypadina Basilewsky, 1984Demetriola Jeannel, 1949Peliocypas Schmidt-Goebel, 1846Subtribe Pericalina Hope, 1838Agonocheila Chaudoir, 1848Allophanes Andrewes, 1939Allophanopsis Louwerens, 1952Amphimenes Bates, 1873Amphimenoides Kirschenhofer, 1999Antimerina Alluaud, 1897Arsinoe Laporte, 1834Auchmerus Andrewes, 1929Bellavalentis Hunting & Yang, 2019Brachichila Chaudoir, 1870Brachyctis Chaudoir, 1870Catascopellus Straneo, 1969Catascopus Kirby, 1825Cheilagona Baehr, 2006Coptodera Dejean, 1825Coptoglossus Chaudoir, 1870Coptoptera Chaudoir, 1837Coptopterella Basilewsky, 1961Coptopterina Basilewsky, 1956Crassagena Baehr, 2006Cylindropectus Lorenz, 1998Dolichoctis Schmidt-Goebel, 1846Dontolobus Basilewsky, 1970Drymatus Motschulsky, 1862Eucheila Dejean, 1829Eurycoleus Chaudoir, 1848Eurydera Laporte, 1831Formosiella Jedlicka, 1951Gidda Andrewes, 1920Glyphodactyla Chaudoir, 1837Holcoderus Chaudoir, 1870Horniulus Jedlicka, 1932Labocephalus Chaudoir, 1848Lelis Chaudoir, 1870Lioptera Chaudoir, 1870Lobodontidius Basilewsky, 1970Lobodontulus Basilewsky, 1970Lobodontus Chaudoir, 1842Madecassina Jeannel, 1949Mascarenhia Alluaud, 1933Menarus Andrewes, 1939Metascopus Basilewsky, 1970Minuphloeus Darlington, 1968Minuthodes Andrewes, 1941Miscelus Klug, 1834Mochtherus Schmidt-Goebel, 1846Mormolyce Hagenbach, 1825Mormolycina Jeannel, 1949Neocoptodera Jeannel, 1949Nycteis Laporte, 1835Oreodicastes Maindron, 1905Oxyodontus Chaudoir, 1870Paradolichoctis Baehr, 2006Pareurydera Jeannel, 1949Pectinitarsus Fairmaire, 1881Pericalus W.S.MacLeay, 1825Philophlaeus Chaudoir, 1844Phloeoxena Chaudoir, 1870Pristacrus Chaudoir, 1870Pseudoplatiella Baehr, 2012Serrimargo Chaudoir, 1870Sfitakantha Andrewes, 1919Sinurus Chaudoir, 1870Stenognathus Chaudoir, 1843Stenotelus Bouchard, 1903Stilboma Andrewes, 1933Stricklandiana Bousquet, 2002Tantillus Chaudoir, 1870Thyreochaetus Basilewsky, 1959Thyreopterus Dejean, 1831Thysanotus Chaudoir, 1848Trichocoptodera Louwerens, 1958Xanthos Kirschenhofer, 2003Xenitenopsis Basilewsky, 1956Xenitenus P\u00e9ringuey, 1896Subtribe Physoderina Chaudoir, 1877Allocota Motschulsky, 1860Anchista Nietner, 1856Dasiosoma Britton, 1937Diamella Shi & Liang, 2013Endynomena Chaudoir, 1873Lachnoderma W.J.MacLeay, 1873Metallanchista Shi & Liang, 2013Orionella Jedlicka, 1964Paraphaea Bates, 1873Physodera Eschscholtz, 1829Subtribe Pseudotrechina Basilewsky, 1984Pseudotrechus Rosenhauer, 1856Subtribe Singilina Jeannel, 1949Singilis Rambur, 1837Subtribe Somotrichina Mateu, 1963Oecornis Britton, 1940Paulianites Jeannel, 1949Pephrica Alluaud, 1936Somotrichus Seidlitz, 1887Subtribe Sugimotoina Habu, 1975Sugimotoa Habu, 1975Subtribe Trichina Basilewsky, 1984Trichis Klug, 1832\u2020Cymindoides Motschulsky, 1856\u2020Lebina Germar, 1813\u2020Miolebidia Zhang; Liu & Shangguan, 1989\u2020Protoscalidion L.Schaufuss, 1888Tribe Odacanthini Laporte, 1834Subtribe Actenonycina Bates, 1871Actenonyx White, 1846Subtribe Homethina Liebherr, 2016Aeolodermus Andrewes, 1929Cyphocoleus Chaudoir, 1877Diplacanthogaster Liebke, 1932Homethes Newman, 1842Quammenis Erwin, 2000Stenocheila Laporte, 1832Subtribe Odacanthina Laporte, 1834Anasis Laporte, 1867Andrewesia Liebke, 1938Arame Andrewes, 1919Archicolliuris Liebke, 1931Asios Liebke, 1933Aulacolius Sloane, 1923Basistichus Sloane, 1917Clarencia Sloane, 1917Colliuris DeGeer, 1774Crassacantha Baehr, 1995Cryptocolliuris Basilewsky, 1955Deipyrodes Bousquet, 2002Dicraspeda Chaudoir, 1863Dobodura Darlington, 1968Erectocolliuris Liebke, 1931Eucolliuris Liebke, 1931Eudalia Laporte, 1867Gestroania Liebke, 1938Giachinoana Baehr, 2003Lachnothorax Motschulsky, 1862Lasiocera Dejean, 1831Mimocolliuris Liebke, 1933Myrmecodemus Sloane, 1923Neoeudalia Baehr, 2005Odacantha Paykull, 1798Ophionea Klug, 1821Polydamasium Liebke, 1938Porocara Sloane, 1917Protocolliuris Liebke, 1931Renneria Baehr, 1999Smeringocera Chaudoir, 1863Stenidia Brull\u00e9, 1834Tricharnhemia Baehr, 2009Subtribe Pentagonicina Bates, 1873Parascopodes Darlington, 1968Pentagonica Schmidt-Goebel, 1846Scopodes Erichson, 1842Tribe Perigonini G.Horn, 1881Diploharpus Chaudoir, 1850Mizotrechus Bates, 1872Perigona Laporte, 1835Ripogenites Basilewsky, 1954Subfamily Licininae Bonelli, 1810[edit]Tribe Chaetogenyini Emden, 1958Camptotoma Reiche, 1843Tribe Chlaeniini Brull\u00e9, 1834Subtribe Callistina Laporte, 1834Callistomimus Chaudoir, 1872Callistus Bonelli, 1810Subtribe Chlaeniina Brull\u00e9, 1834Actodus Alluaud, 1915Chlaenius Bonelli, 1810Eccoptomenus Chaudoir, 1850Ectenognathus Murray, 1858Globulipalpus Sciaky & Facchini, 2019Harpaglossus Motschulsky, 1858Hololeius LaFert\u00e9-S\u00e9nect\u00e8re, 1851Holosoma Semenov, 1889Mirachlaenius Facchini, 2011Parachlaenius Kolbe, 1894Perissostomus Alluaud, 1930Procletodema P\u00e9ringuey, 1898Procletus P\u00e9ringuey, 1896Rhopalomelus Boheman, 1848Sphodroschema Alluaud, 1930Stenoodes Basilewsky, 1953Straneomelus Sciaky & Facchini, 2019Stuhlmannium Kolbe, 1894Viridagonum Lassalle, 2015\u2020Rhopalochlaenius Zhang; Sun & Zhang, 1994Tribe Licinini Bonelli, 1810Subtribe Dicaelina Laporte, 1834Dicaelus Bonelli, 1813Diplocheila Brull\u00e9, 1835Subtribe Dicrochilina Ball, 1992Dicrochile Gu\u00e9rin-M\u00e9neville, 1846Subtribe Lestignathina Ball, 1992Atrotus P\u00e9ringuey, 1896Dilonchus Andrewes, 1936Hormacrus Sloane, 1898Lacordairia Laporte, 1867Lestignathus Erichson, 1842Microferonia Blackburn, 1890Microzargus Sciaky & Facchini, 1997Platylytron W.J.MacLeay, 1873Siagonyx W.J.MacLeay, 1871Zargus Wollaston, 1854Subtribe Licinina Bonelli, 1810Badister Clairville, 1806Colpostoma Semenov, 1889Derostichus Motschulsky, 1860Eurygnathus Wollaston, 1854Eutogeneius Solier, 1849Licinus Latreille, 1802Omestes Andrewes, 1933Physolaesthus Chaudoir, 1850Tribe Oodini LaFert\u00e9-S\u00e9nect\u00e8re, 1851Subtribe Dercylina Sloane, 1923Dercylus Laporte, 1832Subtribe Geobaenina P\u00e9ringuey, 1896Geobaenus Dejean, 1829Subtribe Melanchitonina Jeannel, 1948Dicaelindus W.S.MacLeay, 1825Melanchiton Andrewes, 1940Melanchrous Andrewes, 1940Subtribe Oodina LaFert\u00e9-S\u00e9nect\u00e8re, 1851Acanthoodes Basilewsky, 1953Acutosternus Lecordier & Girard, 1988Adelopomorpha Heller, 1916Anatrichis LeConte, 1853Bamaroodes Gueorguiev, 2014Brachyodes Jeannel, 1949Chaetocrepis Chaudoir, 1857Coptocarpus Chaudoir, 1857Dercylinus Chaudoir, 1883Evolenes LeConte, 1853Holcocoleus Chaudoir, 1883Hoplolenus LaFert\u00e9-S\u00e9nect\u00e8re, 1851Lobatodes Basilewsky, 1956Lonchosternus LaFert\u00e9-S\u00e9nect\u00e8re, 1851Macroprotus Chaudoir, 1878Megaloodes Lesne, 1896Microodes Jeannel, 1949Miltodes Andrewes, 1922Nanodiodes Bousquet, 1996Neoodes Basilewsky, 1953Nothoodes Gueorguiev & Liang, 2020Oodes Bonelli, 1810Oodinus Motschulsky, 1865Orthocerodus Basilewsky, 1946Polychaetus Chaudoir, 1882Prionognathus LaFert\u00e9-S\u00e9nect\u00e8re, 1851Protopidius Basilewsky, 1949Pseudoodes Gueorguiev & Liang, 2020Pseudosphaerodes Jeannel, 1949Simous Chaudoir, 1882Sphaerodinus Jeannel, 1949Sphoerodes Chaudoir, 1883Stenocrepis Chaudoir, 1857Sundaoodes Gueorguiev & Liang, 2020Systolocranius Chaudoir, 1857Thaioodes Gueorguiev, 2014Thryptocerus Chaudoir, 1878Trichopalpoodes B.Gueorguiev & J.Schmidt, 2016Subfamily Loricerinae Bonelli, 1810[edit]Loricera Latreille, 1802Subfamily Melaeninae Csiki, 1933[edit]Cymbionotum Baudi di Selve, 1864Melaenus Dejean, 1831Subfamily Migadopinae Chaudoir, 1861[edit]Tribe Amarotypini Erwin, 1985Amarotypus Bates, 1872Migadopiella Baehr, 2009Tribe Migadopini Chaudoir, 1861Antarctonomus Chaudoir, 1861Aquilex Moret, 1989Calathosoma Jeannel, 1938Calyptogonia Sloane, 1920Decogmus Sloane, 1915Dendromigadops Baehr, 2013Lissopterus G.R.Waterhouse, 1843Loxomerus Chaudoir, 1842Migadopidius Jeannel, 1938Migadops G.R.Waterhouse, 1842Monolobus Solier, 1849Nebriosoma Laporte, 1867Pseudomigadops Jeannel, 1938Rhytidognathus Chaudoir, 1861Stichonotus Sloane, 1910Taenarthrus Broun, 1914Subfamily Nebriinae Laporte, 1834[edit]Tribe Cicindini Csiki, 1927Archaeocindis Kavanaugh & Erwin, 1991Cicindis Bruch, 1908Tribe Nebriini Laporte, 1834Leistus Fr\u00f6lich, 1799Nebria Latreille, 1802Nippononebria Ueno, 1955\u2020Archaeonebria Kavanaugh & J.Schmidt, 2019\u2020Ledouxnebria Deuve, 1998Tribe Notiokasiini Kavanaugh & N\u00e8gre, 1983Notiokasis Kavanaugh & N\u00e8gre, 1983Tribe Notiophilini Motschulsky, 1850Notiophilus Dum\u00e9ril, 1805Tribe Opisthiini Dupuis, 1912Opisthius Kirby, 1837Paropisthius Casey, 1920Tribe Pelophilini Kavanaugh, 1996Pelophila Dejean, 1821Subfamily Nototylinae B\u00e4nninger, 1927[edit]Nototylus Gemminger & Harold, 1868Subfamily Omophroninae Bonelli, 1810[edit]Omophron Latreille, 1802Subfamily Orthogoniinae Schaum, 1857[edit]Tribe Amorphomerini Sloane, 1923Amorphomerus Sloane, 1923Tribe Idiomorphini Bates, 1891Subtribe Idiomorphina Bates, 1891Idiomorphus Chaudoir, 1846Rathymus Dejean, 1831Strigia Brull\u00e9, 1835Subtribe Perochnoristhina Basilewsky, 1973Perochnoristhus Basilewsky, 1973Tribe Orthogoniini Schaum, 1857Subtribe Glyptina  G.Horn, 1881Glyptus Brull\u00e9, 1837Neoglyptus Basilewsky, 1953Subtribe Orthogoniina Schaum, 1857Actenoncus Chaudoir, 1872Anoncopeucus Chaudoir, 1872Hexachaetus Chaudoir, 1872Neoorthogonius Tian & Deuve, 2006Nepalorthogonius Habu, 1979Orthogonius W.S.MacLeay, 1825Subfamily Panagaeinae Bonelli, 1810[edit]Tribe Brachygnathini Basilewsky, 1946Brachygnathus Perty, 1830Tribe Panagaeini Bonelli, 1810Subtribe Bascanina Basilewsky, 1953Bascanus P\u00e9ringuey, 1896Subtribe Panagaeina Bonelli, 1810Adischissus Fedorenko, 2015Calathocosmus Emden, 1928Cintaroa Kasahara, 1989Cintaromorpha H\u00e4ckel & Anichtchenko, 2016Coptia Brull\u00e9, 1835Craspedophorus Hope, 1838Dischissus Bates, 1873Epigraphodes Basilewsky, 1967Epigraphus Chaudoir, 1869Euschizomerus Chaudoir, 1850Geobius Dejean, 1831Micrixys LeConte, 1854Microschemus Andrewes, 1940Panagaeus Latreille, 1802Paregraphus Basilewsky, 1967Peronomerus Schaum, 1854Psecadius Alluaud, 1911Tefflus Leach in Samouelle, 1819Tinoderus Chaudoir, 1879Tinognathus Chaudoir, 1879Trichisia Motschulsky, 1865Tribe Peleciini Chaudoir, 1880Subtribe Agonicina Sloane, 1920Agonica Sloane, 1920Pseudagonica B.Moore, 1960Subtribe Peleciina Chaudoir, 1880Ardistomopsis Straneo & Ball, 1989Disphericus G.R.Waterhouse, 1842Dyschiridium Chaudoir, 1861Eripus Dejean, 1829Pelecium Kirby, 1819Stricteripus Straneo & Ball, 1989Subfamily Patrobinae Kirby, 1837[edit]Tribe Lissopogonini Zamotajlov, 2000Lissopogonus Andrewes, 1923Tribe Patrobini Kirby, 1837Subtribe Deltomerina Chaudoir, 1872Deltomerus Motschulsky, 1850Subtribe Deltomerodina Zamotajlov, 2002Deltomerodes Deuve, 1992Subtribe Patrobina Kirby, 1837Apatrobus Habu & Baba, 1960Apenetretus Kurnakov, 1960Archipatrobus Zamotajlov, 1992Caelopenetretus Zamotajlov & Ito, 2000Chaetapatrobus Lafer, 1996Chinapenetretus Kurnakov, 1963Dimorphopatrobus Casale & Sciaky, 1994Diplous Motschulsky, 1850Grandipenetretus Zamotajlov & Sciaky, 1999Himalopenetretus Zamotajlov, 2002Indopatrobus Zamotajlov & Wrase, 2006Ledouxius Zamotajlov, 1992Minipenetretus Zamotajlov, 2002Minypatrobus Ueno, 1955Naxipenetretus Zamotajlov, 1999Parapatrobus Zamotajlov, 1992Parapenetretus Kurnakov, 1960Patanitretus Zamotajlov, 2002Patrobus Dejean, 1821Penetretus Motschulsky, 1865Platidiolus Chaudoir, 1878Platypatrobus Darlington, 1938Prodiplous Zamotajlov & Sciaky, 2006Qiangopatrobus Zamotajlov, 2002Quasipenetretus Zamotajlov, 2002Tibetopenetretus Zamotajlov & Sciaky, 1999Subfamily Paussinae Latreille, 1806[edit]Tribe Metriini LeConte, 1853Subtribe \u2020Kryzhanovskianina Deuve, 2019\u2020Kryzhanovskiana Kataev & Kirejtshuk, 2019Metrius Eschscholtz, 1829Sinometrius Wrase & J.Schmidt, 2006Tribe Ozaenini Hope, 1838Subtribe Microzaenina Deuve, 2019Microzaena Fairmaire, 1901Subtribe Mystropomina G.Horn, 1881Mystropomus Chaudoir, 1848Subtribe Ozaenina Hope, 1838Crepidozaena Deuve, 2001Entomoantyx Ball & McCleve, 1990Filicerozaena Deuve, 2001Gibbozaena Deuve, 2001Goniotropis G.R.Gray, 1831Inflatozaena Deuve, 2001Mimozaena Deuve, 2001Ozaena Olivier, 1812Pachyteles Perty, 1830Physea Brull\u00e9, 1835Physeomorpha Ogueta, 1963Platycerozaena B\u00e4nninger, 1927Proozaena Deuve, 2001Serratozaena Deuve, 2001Tachypeles Deuve, 2001Tropopsis Solier, 1849Subtribe Pseudozaenina Sloane, 1905Anentmetus Andrewes, 1924Dhanya Andrewes, 1919Eustra Schmidt-Goebel, 1846Itamus Schmidt-Goebel, 1846Pseudozaena Laporte, 1834Sphaerostylus Chaudoir, 1848Tribe Paussini Latreille, 1806Subtribe Carabidomemnina Wasmann, 1928Carabidomemnus Kolbe, 1924Eohomopterus Wasmann, 1919Subtribe Cerapterina Billberg, 1820Arthropterus W.S.MacLeay, 1838Cerapterus Swederus, 1788Megalopaussus Lea, 1906Mesarthropterus Wasmann, 1926Subtribe Heteropaussina Janssens, 1950Heteropaussus J.Thomson, 1860Subtribe Homopterina Wasmann, 1920Homopterus Westwood, 1841Subtribe Paussina Latreille, 1806Allorhopalus Nagel, 2017Ceratoderus Westwood, 1841Euplatyrhopalus Desneux, 1905Lebioderus Westwood, 1838Leleupaussus Luna de Carvalho, 1962Melanospilus Westwood, 1847Paussomorphus Raffray, 1885Paussus Linnaeus, 1775Platyrhopalides Wasmann, 1918Platyrhopalopsis Desneux, 1905Platyrhopalus Westwood, 1833Pterorhopalus Maruyama, 2011Subtribe Pentaplatarthrina Jeannel, 1946Hexaplatarthrus Jeannel, 1955Pentaplatarthrus Westwood, 1833Subtribe \u2020Arthropteritina Luna de Carvalho, 1961\u2020Arthropterites Wasmann, 1926\u2020Cerapterites Wasmann, 1926\u2020Protocerapterus Wasmann, 1926Subtribe \u2020Eopaussina Luna de Carvalho, 1951\u2020Eopaussus Wasmann, 1926Tribe Protopaussini Gestro, 1892Protopaussus Gestro, 1892Subfamily Platyninae Bonelli, 1810[edit]Tribe Omphreini Ganglbauer, 1891Omphreus Dejean, 1828Tribe Platynini Bonelli, 1810Subtribe Enoicina Basilewsky, 1985Abacetodes Straneo, 1939Enoicus P\u00e9ringuey, 1896Subtribe Platynina Bonelli, 1810Achaetocephala Habu, 1975Achaetoprothorax Habu, 1978Aepsera Chaudoir, 1874Agelaea Gen\u00e9, 1839Agonidium Jeannel, 1948Agonobembix Jeannel, 1948Agonoriascus Basilewsky, 1985Agonorites Jeannel, 1951Agonum Bonelli, 1810Altagonum Darlington, 1952Anchomenus Bonelli, 1810Andinocolpodes Perrault, 1991Andrewesius Andrewes, 1939Aparupa Andrewes, 1930Archagonum Basilewsky, 1953Archicolpodes J.Schmidt, 2001Arhytinus Bates, 1889Atranodes Jedlicka, 1953Atranus LeConte, 1847Austroglyptolenus Roig-Ju\u00f1ent, 2003Beckeria Jedlicka, 1931Blackburnia Sharp, 1878Bothrocolpodes Basilewsky, 1985Bruskespar Morvan, 1998Bruskmoal Morvan, 1998Bryanites Valentine, 1987Callidagonum Lorenz, 1998Cardiomera Bassi, 1834Catacolpodes Basilewsky, 1985Celaenagonum Habu, 1978Chaetagonum Burgeon, 1933Chaetosaurus J.Schmidt, 2001Cinctagonum Baehr, 2012Cistelagonum Baehr, 2012Collagonum Baehr, 1995Colpodes W.S.MacLeay, 1825Colpoides Jedlicka, 1931Colpomimus Basilewsky, 1985Colposphodrus Jedlicka, 1953Ctenognathus Fairmaire, 1843Cymenopterus Jeannel, 1948Cyrtopilus Basilewsky, 1985Dalatagonum Fedorenko, 2011Deliaesianum Morvan, 1999Deltocolpodes Morvan, 1992Dendragonum Jeannel, 1948Diacanthostylus Habu, 1978Dicranoncoides Habu, 1978Dicranoncus Chaudoir, 1850Dinocolpodes J.Schmidt, 2001Dirotus W.S.MacLeay, 1825Dister Morvan, 2006Dolichocolpodes Basilewsky, 1985Dyscolus Dejean, 1831Elliptoleus Bates, 1882Epicolpodes Basilewsky, 1985Eucolpodes Jeannel, 1948Euleptus Klug, 1833Euplynes Schmidt-Goebel, 1846Feroniascus Jeannel, 1951Fortagonum Darlington, 1952Galiciotyphlotes Assmann, 1999Gastragonum Darlington, 1952Glaucagonum Habu, 1978Glyptolenoides Perrault, 1991Glyptolenus Bates, 1878Gyrochaetostylus Habu, 1978Habragonum Ueno, 1964Hannaphota Landin, 1955Haplocolpodes Jeannel, 1951Haplopeza Boheman, 1848Helluocolpodes Liebherr, 2005Hemiplatynus Casey, 1920Henvelik Morvan, 1999Herculagonum Baehr, 2002Hikosanoagonum Habu, 1954Idiagonum Darlington, 1952Idiastes Andrewes, 1931Idiocolpodes Basilewsky, 1985Incagonum Liebherr, 1994Iridagonum Darlington, 1952Ischnagonum Kasahara & Sat\u00f4, 1997Jocqueius Basilewsky, 1988Jujiroa Ueno, 1952Kalchdigor Morvan, 1999Kar Morvan, 1998Karnes Morvan, 2010Kaszabellus Jedlicka, 1954Kiwiplatynus Larochelle & Larivi\u00e8re, 2021Klapperichella Jedlicka, 1956Kuceraianum Morvan, 2002Kupeplatynus Larochelle & Larivi\u00e8re, 2021Laevagonum Darlington, 1952Lassalleianum Morvan, 1999Lepcha Andrewes, 1930Leptagonum Kolbe, 1897Leptocolpodes Basilewsky, 1985Letouzeya Bruneau de Mir\u00e9, 1982Liagonum Jeannel, 1948Liamegalonychus Basilewsky, 1963Liocolpodes Basilewsky, 1985Lissagonum Habu, 1978Lithagonum Darlington, 1952Lobocolpodes Basilewsky, 1985Lorostema Motschulsky, 1865Loxocrepis Eschscholtz, 1829Lucicolpodes J.Schmidt, 2000Maculagonum Darlington, 1952Maoriplatynus Larochelle & Larivi\u00e8re, 2021Meleagros Kirschenhofer, 1999Mesocolpodes Basilewsky, 1985Metacolpodes Jeannel, 1948Mexisphodrus Barr, 1965Montagonum Darlington, 1952Mooreagonum Baehr, 2016Morimotoidius Habu, 1954Nebriagonum Darlington, 1952Negreum Habu, 1958Neobatenus Jeannel, 1948Neocolpodes Jeannel, 1948Neodendragonum Basilewsky, 1953Neomegalonychus Jeannel, 1948Nesiocolpodes Jeannel, 1948Nipponagonum Habu, 1978Notagonum Darlington, 1952Notocolpodes Basilewsky, 1985Notoplatynus B.Moore, 1985Nymphagonum Habu, 1978Olisthopus Dejean, 1828Oncostylus Habu, 1978Onotokiba Alluaud, 1927Onycholabis Bates, 1873Onypterygia Dejean, 1831Orophicus Alluaud, 1925Orthotrichus Peyron, 1856Oxygonium Basilewsky, 1951Oxypselaphus Chaudoir, 1843Pachybatenus Basilewsky, 1973Pachyferonia Jeannel, 1951Paracolpodes Basilewsky, 1985Paraliagonum Basilewsky, 1957Paramegalonychus Basilewsky, 1953Paranchodemus Habu, 1978Paranchus Lindroth, 1974Paraplatynus Baehr, 2016Pawgammm Morvan, 1999Platyagonum Habu, 1978Platynus Bonelli, 1810Plaumannium Liebke, 1939Plicagonum Darlington, 1952Ponapagonum Darlington, 1970Potamagonum Darlington, 1952Praepristus Kirschenhofer, 1999Promecoptera Dejean, 1831Promegalonychus Basilewsky, 1953Prophenorites Basilewsky, 1985Prosphodrus Britton, 1959Protocolpodes Basilewsky, 1985Pseudanchomenus Tarnier, 1860Pseudobatenus Basilewsky, 1951Pseudomegalonychus Basilewsky, 1950Rhadine LeConte, 1846Rupa Jedlicka, 1935Scotagonum Habu, 1978Sericoda Kirby, 1837Shibataia Habu, 1978Sinocolpodes J.Schmidt, 2001Skoeda Morvan, 1995Skorlagad Morvan, 1999Skouedhirraad Morvan, 1999Sophroferonia Alluaud, 1933Speagonum B.Moore, 1977Speocolpodes Barr, 1974Speokokosia Alluaud, 1932Sperkanhir Morvan, 2010Stenocnemus Mannerheim, 1837Straneoa Basilewsky, 1953Syletor Tschitscherine, 1899Takasagoagonum Habu, 1977Tanystoma Motschulsky, 1845Tarsagonum Darlington, 1952Tetraleucus Casey, 1920Tostkar Morvan, 1998Trogloagonum Casale, 1982Tuiplatynus Larochelle & Larivi\u00e8re, 2021Vitagonum B.Moore, 1999Vulcanophilus Heller, 1898Xestagonum Habu, 1978\u2020Praeanchodemus J.Schmidt; G\u00f6pel & Will, 2017Tribe Sphodrini Laporte, 1834Subtribe Atranopsina Baehr, 1982Amaroschema Jeannel, 1943Amphimasoreus Piochard de la Br\u00fblerie, 1875Atranopsis Baehr, 1982Broter Andrewes, 1923Gomerina Bolivar y Pieltain, 1940Paraeutrichopus Mateu, 1954Platyderus Stephens, 1827Pseudomyas Uyttenboogaart, 1929Pseudoplatyderus Bolivar y Pieltain, 1940Subtribe Calathina Laporte, 1834Calathus Bonelli, 1810Thermoscelis Putzeys, 1873Subtribe Dolichina Brull\u00e9, 1834Acalathus Semenov, 1889Anchomenidius Heyden, 1880Casaleius Sciaky & Wrase, 1998Dolichus Bonelli, 1810Doliodactyla Sciaky & Wrase, 1998Morphodactyla Semenov, 1889Xestopus Andrewes, 1937Subtribe Pristosiina Lindroth, 1956Pristosia Motschulsky, 1865Subtribe Sphodrina Laporte, 1834Calathidius Putzeys, 1873Cephalosdrophus Lassalle & Marcilhac, 1999Eosphodrus Casale, 1988Eremosphodrus Semenov, 1909Himalosphodrus Casale, 1988Hystricosphodrus Casale & Giachino, 2004Ifridytes Deuve & Queinnec, 1994Laemostenus Bonelli, 1810Licinopsis Bedel, 1899Miquihuana Barr, 1982Pseudotaphoxenus L.Schaufuss, 1865Reflexisphodrus Casale, 1988Sphodropsis Seidlitz, 1887Sphodrus Clairville, 1806Stenolepta Semenov, 1889Taphoxenus Motschulsky, 1850Subtribe Synuchina Lindroth, 1956Nipponosynuchus Morita, 1998Parabroscus Lindroth, 1956Synuchus Gyllenhal, 1810Trephionus Bates, 1883Subfamily Promecognathinae LeConte, 1853[edit]Tribe Axinidiini Basilewsky, 1963Axinidium Sturm, 1843Holaxinidium Basilewsky, 1963Metaxinidium Basilewsky, 1963Paraxinidium Basilewsky, 1963Tribe Dalyatini Mateu, 2002Dalyat Mateu, 2002Tribe Promecognathini LeConte, 1853Promecognathus Chaudoir, 1846Tribe \u2020Palaeoaxinidiini McKay, 1991\u2020Palaeoaxinidium McKay, 1991Subfamily Pseudomorphinae Hope, 1838[edit]Adelotopus Hope, 1836Cainogenion Notman, 1925Cryptocephalomorpha Ritsema, 1875Guyanemorpha Erwin, 2013Manumorpha Erwin & Geraci, 2008Notopseudomorpha Baehr, 1997Paussotropus C.O.Waterhouse, 1877Pseudomorpha Kirby, 1823Samiriamorpha Erwin & Geraci, 2008Sphallomorpha Westwood, 1840Tuxtlamorpha Erwin & Geraci, 2008Yasunimorpha Erwin & Geraci, 2008Subfamily Psydrinae LeConte, 1853[edit]Tribe Gehringiini Darlington, 1933Subtribe Gehringiina Darlington, 1933Gehringia Darlington, 1933Subtribe Helenaeina Deuve, 2007Afrogehringia Baehr; Sch\u00fcle & Lorenz, 2009Helenaea Schatzmayr & Koch, 1934Tribe Moriomorphini Sloane, 1890Subtribe Amblytelina Blackburn, 1892Amblytelus Erichson, 1842Dystrichothorax Blackburn, 1892Epelyx Blackburn, 1892Mecyclothorax Sharp, 1903Meonis Laporte, 1867Paratrichothorax Baehr, 2004Pseudamblytelus Baehr, 2004Raphetis B.Moore, 1963Selenochilus Chaudoir, 1878Trichamblytelus Baehr, 2004Subtribe Moriomorphina Sloane, 1890Celanida Laporte, 1867Melisodera Westwood, 1835Molopsida White, 1846Moriodema Laporte, 1867Moriomorpha Laporte, 1867Neonomius B.Moore, 1963Pterogmus Sloane, 1920Rhaebolestes Sloane, 1903Rossjoycea Liebherr, 2011Sitaphe B.Moore, 1963Spherita Liebherr, 2020Tarastethus Sharp, 1883Teraphis Laporte, 1867Theprisa B.Moore, 1963Trephisa B.Moore, 1963Trichopsida Larochelle & Larivi\u00e8re, 2013Subtribe Tropopterina Sloane, 1898Pharetis Liebherr, 2020Tropopterus Solier, 1849Tribe Psydrini LeConte, 1853Laccocenus Sloane, 1890Nomius Laporte, 1835Psydrus LeConte, 1846Subfamily Pterostichinae Bonelli, 1810[edit]Tribe Chaetodactylini Tschitscherine, 1903Chaetodactyla Tschitscherine, 1897Tribe Cnemalobini Germain, 1911Cnemalobus Gu\u00e9rin-M\u00e9neville, 1838Tribe Cratocerini Lacordaire, 1854Subtribe Catapieseina Bates, 1882Catapiesis Solier, 1835Homalomorpha Brull\u00e9, 1837Subtribe Cratocerina Lacordaire, 1854Brachidius Chaudoir, 1852Cratocerus Dejean, 1829Oxyglychus Straneo, 1938Subtribe Drimostomatina Chaudoir, 1873Abacaelostus Straneo, 1952Andrewesinulus Straneo, 1938Apsidocnemus Alluaud, 1936Barylaus Liebherr, 1985Caecocaelus Straneo, 1949Caelostomus W.S.MacLeay, 1825Camptogenys Tschitscherine, 1899Capabatus Csiki, 1930Crenulostrigus Straneo, 1942Cyrtolaus Bates, 1882Dactyleurys Tschitscherine, 1899Dactylinius Straneo, 1941Diachipteryx Alluaud, 1925Diceromerus Chaudoir, 1873Dromistomus Jeannel, 1948Drymonaxus Straneo, 1941Feostoma Straneo, 1941Hemitelestus Alluaud, 1895Hoplizomenus Chaudoir, 1873Leleuporites Straneo, 1960Madapelmus Dajoz, 1985Monodryxus Straneo, 1942Pachycaecus Straneo, 1971Pachyroxochus Straneo, 1942Platyxythrius Lorenz, 1998Stegazopteryx Will, 2004Stomonaxellus Tschitscherine, 1901Strigomerodes Straneo, 1939Strigomerus Chaudoir, 1873Trichillinus Straneo, 1938Tribe Microcheilini Jeannel, 1948Microcheila Brull\u00e9, 1835Tribe Morionini Brull\u00e9, 1837Buderes Murray, 1857Hyperectenus Alluaud, 1935Hyperion Laporte, 1834Megamorio Chaudoir, 1880Morion Latreille, 1810Morionidius Chaudoir, 1880Moriosomus Motschulsky, 1855Platynodes Westwood, 1847Stereostoma Murray, 1857Tribe Pterostichini Bonelli, 1810Subtribe Abacetina Chaudoir, 1873Abacaecus Allegro & Giachino, 2020Abacetus Dejean, 1828Aristopus LaFert\u00e9-S\u00e9nect\u00e8re, 1853Celioinkosa Straneo, 1951Chlaeminus Motschulsky, 1865Colpodichius Straneo, 1952Cosmodiscus Sloane, 1907Cyrtomoscelis Chaudoir, 1874Distrigidius Jeannel, 1948Ecnomolaus Bates, 1892Haptoderidius Straneo, 1975Haptoderodes Straneo, 1986Holconotus Schmidt-Goebel, 1846Inkosa P\u00e9ringuey, 1926Mateuellus Deuve, 1990Metabacetus Bates, 1892Metaxellus Straneo, 1960Metaxys Chaudoir, 1857Novillidius Straneo, 1941Oodinkosa Straneo, 1939Ophonichius Straneo, 1942Pediomorphus Chaudoir, 1878Pioprosopus Tschitscherine, 1899Pollicobius Vinson, 1939Prostalomus Basilewsky, 1950Pseudabacetus Burgeon, 1935Pterostillichus Straneo, 1949Rhagadillius Straneo, 1951Tiferonia Darlington, 1962Trachelocyphoides Straneo, 1942Trachelocyphus Tschitscherine, 1900Subtribe Euchroina Chaudoir, 1874Abacillius Straneo, 1949Abacillodes Straneo, 1988Abaris Dejean, 1831Apsaustodon Tschitscherine, 1901Argutoridius Chaudoir, 1876Blennidus Motschulsky, 1866Blenniventer Straneo, 1986Bothynoproctus Tschitscherine, 1900Cedrorum Borges & A.Serrano, 1993Cephalostichus Straneo, 1977Cynthidia Chaudoir, 1874Euchroa Brull\u00e9, 1835Eumara Tschitscherine, 1901Gastrogmus Sloane, 1915Haplobothynus Tschitscherine, 1901Litarthrum Sloane, 1915Lobobrachus Sharp, 1885Marsyas Putzeys, 1845Meropalpus Tschitscherine, 1900Microcephalus Dejean, 1828Neotalus Will, 2002Oribazus Chaudoir, 1874Orthomus Chaudoir, 1838Pachythecus Chaudoir, 1874Paniestichus Will, 2011Parorthomus B.Gueorguiev; Wrase & Farkac, 2014Phaenaulax Tschitscherine, 1898Prosopogmus Chaudoir, 1865Pseudabarys Chaudoir, 1874Setalimorphus Sloane, 1895Setalis Laporte, 1867Simodontus Chaudoir, 1843Trirammatus Chaudoir, 1835Subtribe Loxandrina Erwin & Sims, 1984Cerabilia Laporte, 1867Oxycrepis Reiche, 1843Zeodera Laporte, 1867Subtribe Metiina Straneo, 1951Abropus G.R.Waterhouse, 1842Antarctiola Straneo, 1977Feroniola Tschitscherine, 1900Kuschelinus Straneo, 1963Metius Curtis, 1839Subtribe Pterostichina Bonelli, 1810Abacidus LeConte, 1863Abacoleptus Fauvel, 1903Abacomorphus Chaudoir, 1878Abacophrastus Will, 2011Abacops Tschitscherine, 1902Abax Bonelli, 1810Allotriopus Bates, 1882Analoma Darlington, 1971Apophylon B.Gueorguiev & Sciaky, 2015Aristochroa Tschitscherine, 1898Aristochroodes Marcilhac, 1993Aulacopodus Britton, 1940Basilewskya Straneo, 1948Camptoscelis Dejean, 1828Castelnaudia Tschitscherine, 1891Catadromus W.S.MacLeay, 1825Chaetauchenium Tschitscherine, 1900Chalcochrous Chaudoir, 1838Cophosomorpha Tschitscherine, 1891Cratoferonia Tschitscherine, 1902Cratogaster Blanchard, 1843Cuneipectus Sloane, 1907Cyclotrachelus Chaudoir, 1838Cyrtoderus Hope, 1842Darodilia Laporte, 1867Delinius Westwood, 1864Eucamptognathus Chaudoir, 1837Eudromus Klug, 1835Euryabax Fauvel, 1903Euryaptus Bates, 1892Eurypercus Jeannel, 1948Eurystomis Chaudoir, 1878Gastrellarius Casey, 1918Gourlayia Britton, 1964Harpostomus Chaudoir, 1856Henrotiochoromus Busulini, 1958Henrotius Jeannel, 1953Holcaspis Chaudoir, 1865Hybothecus Chaudoir, 1874Leiolesticus Roux; Lassalle & Dubault, 2016Lesticus Dejean, 1828Licentius Jedlicka, 1939Liopasa Tschitscherine, 1901Lophoglossus LeConte, 1853Loxogenius Sloane, 1907Mecynognathus W.J.MacLeay, 1873Megadromus Motschulsky, 1866Molopidius Jeannel, 1942Molopinus Jeannel, 1948Molops Bonelli, 1810Myas Sturm, 1826Neoferonia Britton, 1940Nesites Andrewes, 1939Nirmala Andrewes, 1930Notabax B.Moore, 1976Notolestus Sloane, 1894Notonomus Chaudoir, 1865Nurus Motschulsky, 1866Oberthueria Vuillet, 1911Ogmophora Tschitscherine, 1896Onawea Johns, 2007Oscadytes Lagar Mascaro, 1975Pachymorphus Chaudoir, 1838Paranurus Tschitscherine, 1901Pareuryaptus Dubault; Lassalle & Roux, 2008Parhypates Motschulsky, 1866Pedius Motschulsky, 1850Percolaus Bates, 1882Percus Bonelli, 1810Peyrieraselus Deuve, 1981Piesmus LeConte, 1846Platycoelus Blanchard, 1843Platysmodes Fauvel, 1903Plocamostethus Britton, 1940Poecilinus Jeannel, 1948Poecilus Bonelli, 1810Pseudoceneus Tschitscherine, 1891Pterostichus Bonelli, 1810Rhabdotus Chaudoir, 1865Rhytiferonia Darlington, 1962Rhytisternus Chaudoir, 1865Sarticus Motschulsky, 1866Secatophus Laporte, 1867Setalidius Chaudoir, 1878Speluncarius Reitter, 1886Speomolops Patrizi, 1955Sphodrosomus Perroud, 1864Stenochoromus L.Miller, 1866Stereocerus Kirby, 1837Sthenocranion Tschitscherine, 1899Stomis Clairville, 1806Straneostichus Sciaky, 1994Styracoderus Chaudoir, 1874Tanythrix Schaum, 1858Taphoxomimus Straneo, 1975Tapinopterus Schaum, 1858Teratotarsa Tschitscherine, 1893Teropha Laporte, 1867Trichosternus Chaudoir, 1865Trigonaptus Fedorenko, 2020Trigonotoma Dejean, 1828Troglorites Jeannel, 1919Tropidocerus Chaudoir, 1878Typhlochoromus Moczarski, 1913Wahlbergiana Bousquet, 2002Xenion Tschitscherine, 1902Zariquieya Jeannel, 1924Zeopoecilus Sharp, 1886Tribe Zabrini Bonelli, 1810Subtribe Amarina Zimmermann, 1832Amara Bonelli, 1810Pseudamara Lindroth, 1968Subtribe Zabrina Bonelli, 1810Zabrus Clairville, 1806Subfamily Rhysodinae Laporte, 1840[edit]Tribe Clinidiini R.T. & J.R.Bell, 1978Clinidium Kirby, 1830Grouvellina R.T. & J.R.Bell, 1978Rhyzodiastes Fairmaire, 1895Tribe Dhysorini R.T. & J.R.Bell, 1978Dhysores Grouvelle, 1903Neodhysores R.T. & J.R.Bell, 1978Tangarona R.T. & J.R.Bell, 1982Tribe Leoglymmiini R.T. & J.R.Bell, 1978Leoglymmius R.T. & J.R.Bell, 1978Tribe Medisorini R.T. & J.R.Bell, 1987Medisores R.T. & J.R.Bell, 1987Tribe Omoglymmiini R.T. & J.R.Bell, 1978Arrowina R.T. & J.R.Bell, 1978Omoglymmius Ganglbauer, 1891Plesioglymmius R.T. & J.R.Bell, 1978Shyrodes Grouvelle, 1903Srimara R.T. & J.R.Bell, 1978Xhosores R.T. & J.R.Bell, 1978Yamatosa R.T. & J.R.Bell, 1979Tribe Rhysodini Laporte, 1840Kaveinga R.T. & J.R.Bell, 1978Kupeus R.T. & J.R.Bell, 1982Rhysodes Germar, 1822Tribe Sloanoglymmiini R.T. & J.R.Bell, 1991Sloanoglymmius R.T. & J.R.Bell, 1991Subfamily Scaritinae Bonelli, 1810[edit]Tribe Clivinini Rafinesque, 1815Subtribe Androzelmina R.T.Bell, 1998Androzelma Dostal, 1993Subtribe Ardistomina Putzeys, 1867Ardistomis Putzeys, 1846Aspidoglossa Putzeys, 1846Kearophus Dajoz, 2004Semiardistomis Kult, 1950Subtribe Clivinina Rafinesque, 1815Afrosyleter Basilewsky, 1959Ancus Putzeys, 1867Basilewskyana Kult, 1959Brachypelus Putzeys, 1867Cameroniola Baehr, 2000Climax Putzeys, 1861Clivina Latreille, 1802Clivinarchus Sloane, 1896Cryptomma Putzeys, 1846Lachenus Putzeys, 1846Leleuporella Basilewsky, 1956Nannoryctes Baehr, 2000Nyctosyles Putzeys, 1867Orictites Andrewes, 1931Platysphyrus Sloane, 1905Psilidius Jeannel, 1957Pyramoides Bousquet, 2002Rhysocara Sloane, 1916Rubidiclivina Baehr, 2015Rugiluclivina Balkenohl, 1996Semiclivina Kult, 1947Sinesetosa Balkenohl, 1996Syleter Andrewes, 1941Thliboclivina Kult, 1959Trilophidius Jeannel, 1957Trilophus Andrewes, 1927Trogloclivina Deuve, 2003Whiteheadiana Perrault, 1994Subtribe Forcipatorina B\u00e4nninger, 1938Camptidius Putzeys, 1867Camptodontus Dejean, 1826Forcipator Maindron, 1904Kultianella Perrault, 1994Mesus Chevrolat, 1858Obadius Burmeister, 1875Oxygnathopsis Louwerens, 1953Oxygnathus Dejean, 1826Scolyptus Putzeys, 1861Stratiotes Putzeys, 1846Subtribe Reicheiina Jeannel, 1957Alpiodytes Jeannel, 1957Asioreicheia Bulirsch & Magrini, 2014Catalanodytes Sciaky, 1989Dalmatoreicheia Magrini & Bulirsch, 2005Dimorphoreicheia Magrini; Fancello & Leo, 2002Galicioreicheia Felix & Bulirsch, 2015Guiodytes Tian, 2013Gymnetoreicheia Magrini; Fancello & Casale, 2019Iberodytes Jeannel, 1949Ichnusodytes Magrini; Fancello & Onnis, 2019Italodytes G.M\u00fcller, 1938Kenyoreicheia Bulirsch & Magrini, 2007Laoreicheia Balkenohl, 2005Madagascareicheia Magrini & Bulirsch, 2009Malagasyreicheia Magrini; Bulirsch & Fancello, 2021Orientoreicheia Bulirsch & Hurka, 1994Oxydrepanus Putzeys, 1867Parareicheia Jeannel, 1957Reicheadella Reitter, 1913Reicheia Saulcy, 1862Reicheidius Jeannel, 1957Sikelioreicheia Magrini & Fancello, 2019Spelaeodytes L.Miller, 1863Typhloreicheia Holdhaus, 1924Subtribe Schizogeniina Dostal, 2017Baehrogenius Dostal, 2017Coryza Putzeys, 1867Halocoryza Alluaud, 1919Lophocoryza Alluaud, 1941Paracoryza Basilewsky, 1952Psammocoryza Hogan, 2006Schizogenius Putzeys, 1846Subtribe Sparostesina Dostal, 2017Bohemaniella Bousquet, 2002Pseudoclivina Kult, 1947Sparostes Putzeys, 1867Tribe Corintascarini Basilewsky, 1973Corintascaris Basilewsky, 1952Tribe Dyschiriini Kolbe, 1880Akephorus LeConte, 1852Caledyschirius Bulirsch, 2010Clivinopsis Bedel, 1895Cribrodyschirius Bruneau de Mir\u00e9, 1952Dyschirius Bonelli, 1810Neodyschirius Kult, 1954Reicheiodes Ganglbauer, 1891Setodyschirius Fedorenko, 1996Striganoviella Fedorenko, 2012Torretassoa Schatzmayr & Koch, 1933\u2020Dyschiriomimus Iablokoff-Khnzorian, 1960Tribe Salcediini Alluaud, 1930Subtribe Salcediina Alluaud, 1930Salcedia Fairmaire, 1899Subtribe Solenogenyina R.T.Bell, 1998Holoprizus Putzeys, 1867Solenogenys Westwood, 1859Tribe Scaritini Bonelli, 1810Subtribe Carenina W.J.MacLeay, 1887Carenidium Chaudoir, 1868Carenum Bonelli, 1813Epilectus Blackburn, 1888Euryscaphus W.J.MacLeay, 1865Laccopterum W.J.MacLeay, 1878Monocentrum Chaudoir, 1868Mouhotia Laporte, 1862Neocarenum Laporte, 1867Neoscaphus Sloane, 1888Philoscaphus W.J.MacLeay, 1871Scaraphites Westwood, 1842Trichocarenum Blackburn, 1892Subtribe Pasimachina Putzeys, 1867Pasimachus Bonelli, 1813Subtribe Scapterina Putzeys, 1867Acanthoscelis Dejean, 1825Oxylobus Chaudoir, 1855Parathlibops Basilewsky, 1958Passalidius Chaudoir, 1863Scapterus Dejean, 1826Steganomma W.J.MacLeay, 1887Thlibops Putzeys, 1867Subtribe Scaritina Bonelli, 1810Anomophaenus Fauvel, 1882Antilliscaris Lorenz, 1998Baenningeria Reichardt, 1976Coptolobus Chaudoir, 1857Crepidopterus Chaudoir, 1855Cryptoscaphus Chaudoir, 1855Dinoscaris Alluaud, 1902Distichus Motschulsky, 1858Dyscaris B\u00e4nninger, 1940Dyscherinus Jeannel, 1955Dyscherus Chaudoir, 1855Geoscaptus Chaudoir, 1855Glyptogrus Bates, 1875Gnaphon Andrewes, 1920Haplogaster Chaudoir, 1879Haplotrachelus Chaudoir, 1855Macromorphus Chaudoir, 1857Madascaris B\u00e4nninger, 1938Mamboicus Bates, 1886Mecynoscaris Alluaud, 1930Menigius Chaudoir, 1879Neochryopus B\u00e4nninger, 1932Ochyropus Schi\u00f6dte, 1847Pachyodontus Chaudoir, 1879Paradyscherus Basilewsky, 1971Pilades Heyne, 1895Prodyscherodes Jeannel, 1955Prodyscherus Jeannel, 1946Scarites Fabricius, 1775Storthodontus Chaudoir, 1855Tapinoscaris Jeannel, 1946Tibioscarites B\u00e4nninger, 1929Tonkinoscaris B\u00e4nninger, 1956Typhloscaris Kuntzen, 1914Subfamily Siagoninae Bonelli, 1813[edit]Tribe Enceladini G.Horn, 1881Enceladus Bonelli, 1813Tribe Siagonini Bonelli, 1813Luperca Laporte, 1840Siagona Latreille, 1804Subfamily Trechinae Bonelli, 1810[edit]Tribe Bembidarenini Maddison et al., 2019Andinodontis Erwin & Toledano, 2010Argentinatachoides Sallanave; Erwin & Roig-Ju\u00f1ent, 2008Bembidarenas Erwin, 1972Tasmanitachoides Erwin, 1972Tribe Bembidiini Stephens, 1827Subtribe Anillina Jeannel, 1937Afranillus Giachino, 2015Afrodipnus Giachino, 2015Angustanillus Baehr & Main, 2016Anillaspis Casey, 1918Anillinus Casey, 1918Anillodes Jeannel, 1963Anillopsidius Coiffait, 1969Anillopsis Jeannel, 1937Anillotarsus Mateu, 1980Anillus Jacquelin du Val, 1851Argiloborus Jeannel, 1937Austranillus Giachino, 2005Bafutyphlus Bruneau de Mir\u00e9, 1986Bhutanillus M.E.Schmid, 1975Bulirschia Giachino, 2008Bylibaraphanus Giachino; Eberhard & Perina, 2021Caeconannus Jeannel, 1963Caecoparvus Jeannel, 1937Carayonites Bruneau de Mir\u00e9, 1986Cryptocharidius M.Etonti & Mateu, 1992Cryptorites Jeannel, 1950Dicropterus Ehlers, 1883Elgonotyphlus Sciaky & Zaballos, 1993Erwinanillus Giachino; Eberhard & Perina, 2021Externanillus Baehr & Main, 2016Geocharidius Jeannel, 1963Geocharis Ehlers, 1883Gilesdytes Giachino; Eberhard & Perina, 2021Gracilanillus Baehr & Main, 2016Gregorydytes Giachino; Eberhard & Perina, 2021Hesperanillus Baehr & Main, 2016Honduranillus Zaballos, 1997Hygranillus B.Moore, 1980Hypodipnites Jeannel, 1963Hypotyphlus Jeannel, 1937Iason Giachino & Vailati, 2011Iberanillus Espa\u00f1ol, 1971Illaphanus W.J.MacLeay, 1865Kimberleytyphlus Giachino; Eberhard & Perina, 2021Leleupanillus Basilewsky, 1976Magnanillus Baehr, 2017Malagasydipnus Giachino, 2008Malagasytyphlus Giachino, 2008Megastylulus Giachino & Sciaky, 2003Mexanillus Vigna Taglianti, 1973Microdipnites Jeannel, 1957Microdipnodes Basilewsky, 1960Microdipnus Jeannel, 1937Microtyphlus Linder, 1863Mystroceridius Reichardt, 1972Neodipnus Jeannel, 1957Neoillaphanus Giachino; Eberhard & Perina, 2021Nesamblyops Jeannel, 1937Nothanillus Jeannel, 1962Orthotyphlus Mateu & Zaballos, 1998Paranillopsis Cicchino & Roig-Ju\u00f1ent, 2001Paranillus Jeannel, 1949Parillaphanus Baehr, 2018Parvocaecus Coiffait, 1956Pelocharis Jeannel, 1960Pelodiaetodes B.Moore, 1980Pelodiaetus Jeannel, 1937Pelonomites Jeannel, 1963Perucharidius Mateu & M.Etonti, 2002Pilbaradytes Giachino; Eberhard & Perina, 2021Pilbaranillus Baehr & Main, 2016Pilbaraphanus Giachino; Eberhard & Perina, 2021Prioniomus Jeannel, 1937Pseudanillus Bedel, 1896Pseudillaphanus Giachino, 2005Rhegmatobius Jeannel, 1937Scotodipnus Schaum, 1860Selenodipnus Jeannel, 1963Serranillus Barr, 1995Serratotyphlus Giachino, 2008Stylulus L.Schaufuss, 1882Tasmanillus Giachino, 2005Typhlocharis Dieck, 1869Typhlonesiotes Jeannel, 1937Winklerites Jeannel, 1937Zapotecanillus Sokolov, 2013Zeanillus Jeannel, 1937Zoianillus Sciaky, 1994Subtribe Bembidiina Stephens, 1827Amerizus Chaudoir, 1868Asaphidion Gozis, 1886Bembidion Latreille, 1802Caecidium Ueno, 1971Lionepha Casey, 1918Ocys Stephens, 1828Orzolina Machado, 1987Sinechostictus Motschulsky, 1864Subtribe Horologionina Jeannel, 1949Horologion Valentine, 1932Subtribe Lovriciina Giachino; B.Gueorguiev & Vailati, 2011Lovricia Pretner, 1979Neolovricia Lakota; Jalzic & J.Moravec, 2009Paralovricia Giachino; B.Gueorguiev & Vailati, 2011Subtribe Tachyina Motschulsky, 1862Aenigmatachys Baehr, 2020Afrotachys Basilewsky, 1958Anomotachys Jeannel, 1946Costitachys Erwin, 1974Elaphropus Motschulsky, 1839Kiwitachys Larochelle & Larivi\u00e8re, 2007Liotachys Bates, 1871Lymnastis Motschulsky, 1862Meotachys Erwin, 1974Micratopus Casey, 1914Moirainpa Erwin, 1984Nothoderis Boyd & Erwin, 2016Pericompsus LeConte, 1852Polyderidius Jeannel, 1962Porotachys Netolitzky, 1914Setitachys Baehr, 2016Stigmatachys Boyd & Erwin, 2016Straneoites Basilewsky, 1947Tachys Dejean, 1821Tachysbembix Erwin, 2004Tachyta Kirby, 1837Tachyxysta Boyd & Erwin, 2016Subtribe Xystosomina Erwin, 1994Erwiniana Paulsen & A.Smith, 2003Geballusa Erwin, 1994Gouleta Erwin, 1994Inpa Erwin, 1978Mioptachys Bates, 1882Philipis Erwin, 1994Xystosomus Schaum, 1863Tribe Pogonini Laporte, 1834Bedeliolus Semenov, 1900Cardiaderus Dejean, 1828Diodercarus Lutshnik, 1931Diplochaetus Chaudoir, 1872Ochtozetus Chaudoir, 1872Olegius Komarov, 1996Pogonistes Chaudoir, 1872Pogonopsis Bedel, 1898Pogonus Dejean, 1821Sirdenus Dejean, 1828Syrdenoidius Baehr & Hudson, 2001Thalassotrechus Van Dyke, 1918Tribe Sinozolini Deuve, 1997Chaltenia Roig-Ju\u00f1ent & Cicchino, 2001Phrypeus Casey, 1924Sinozolus Deuve, 1997Tribe Trechini Bonelli, 1810Subtribe Aepina Fowler, 1887Aepopsis Jeannel, 1922Aepus Leach in Samouelle, 1819Kenodactylus Broun, 1909Kiwitrechus Larochelle & Larivi\u00e8re, 2007Maoritrechus Brookes, 1932Neanops Britton, 1962Oarotrechus Townsend, 2010Temnostega Enderlein, 1905Thalassobius Solier, 1849Waiputrechus Townsend, 2010Subtribe Perileptina Sloane, 1903Apoplotrechus Alluaud, 1915Neoblemus Jeannel, 1923Perileptus Schaum, 1860Subtribe Trechina Bonelli, 1810Accoella Ueno, 1990Acheroniotes Lohaj & Lakota, 2010Adriaphaenops Noesske, 1928Aepiblemus Belousov & Kabak, 1993Agonotrechus Jeannel, 1923Agostinia Jeannel, 1928Alanorites Belousov, 1998Albanotrechus Casale & V.B.Gueorguiev, 1994Allegrettia Jeannel, 1928Allotrechiama Ueno, 1970Ameroduvalius Valentine, 1952Anchotrechus Jeannel, 1927Andinorites Mateu & Belles, 1980Andinotrechus Mateu, 1981Anillidius Jeannel, 1928Anophthalmus Sturm, 1844Aphaenopidius G.M\u00fcller, 1913Aphaenops Bonvouloir, 1862Aphaenopsis G.M\u00fcller, 1913Apocimmerites Belousov, 1998Apoduvalius Jeannel, 1953Aputrechisibus Trezzi, 2007Arctaphaenops Meixner, 1925Aspidaphaenops Ueno, 2006Austrotrechus B.Moore, 1972Awatrechus Ueno, 1955Balazucellus Deuve, 2001Bathytrechus Ueno, 2005Beronaphaenops B.V.Gueorguiev, 2012Birmaphaenops Deuve, 2017Blemus Dejean, 1821Boldoriella Jeannel, 1928Boreaphaenops Ueno, 2002Bothynotrechus B.Moore, 1972Casaleaphaenops Tian; Huang & Ma, 2021Cathaiaphaenops Deuve, 1996Caucasaphaenops Belousov, 1999Caucasorites Belousov & Zamotajlov, 1997Chaetoduvalius Jeannel, 1928Chaetotrechiama Ueno, 1982Chiapadytes Vigna Taglianti, 1977Chu Tian & He, 2020Cimbrodytes Piva, 2021Cimmerites Jeannel, 1928Cimmeritodes Deuve, 1996Ckacus Tian & Huang, 2021Columbitrechus Mateu, 1982Coreoblemus Ueno, 1969Croatotrechus Casale & Jalzic, 1999Dabatrechus Ueno, 2004Dactylotrechus Belousov & Kabak, 2003Daiconotrechus Ueno, 1971Dalmataphaenops Monguzzi, 1993Darlingtonea Valentine, 1952Derossiella Queinnec, 2008Deuveaphaenops Tian & Huang, 2017Dianotrechus Tian, 2016Doderotrechus Vigna Taglianti, 1968Dongoblemus Deuve & Tian, 2016Dongodytes Deuve, 1993Dracotrechus Ueno, 2010Duvalioblemus Deuve, 1995Duvaliomimus Jeannel, 1928Duvaliopsis Jeannel, 1928Duvalius Delarouz\u00e9e, 1859Eocnides Jeannel, 1954Epaphiopsis Ueno, 1953Epaphiotrechus Deuve & Kavanaugh, 2016Epaphius Leach in Samouelle, 1819Erebotrechus Britton, 1964Escolatrechus Mateu, 2003Eutrechopsis B.Moore, 1972Eutrechus B.Moore, 1972Geotrechus Jeannel, 1919Giraffaphaenops Deuve, 2002Glabroduvalius Vrbica; Curcic; Antic & Curcic, 2013Goedetrechus B.Moore, 1972Gotoblemus Ueno, 1970Graciliblemus Deuve & Tian, 2016Guatemalatrechus Trezzi, 2003Guiaphaenops Deuve, 2002Guizhaphaenops Vigna Taglianti, 1997Guizhaphaenopsoides Tian; Huang & Li, 2021Gulaphaenops Ueno, 1987Himalaphaenops Ueno, 1980Himalotrechus Belousov; Kabak & J.Schmidt, 2019Homaloderodes Jeannel, 1962Huoyanodytes Tian & Huang, 2016Iberotrechus Jeannel, 1920Iga Ueno, 1953Incatrechus Mateu & Belles, 1982Inotrechus Dolzhanski & Ljovushkin, 1989Ishidatrechus Ueno, 1956Ishikawatrechus Habu, 1950Italaphaenops Ghidini, 1964Iyotrechus Ueno & Naito, 2009Jalzicaphaenops Lohaj & Lakota, 2010Jeannelius Kurnakov, 1959Jiangxiaphaenops Ueno & Clarke, 2007Jiulongotrechus Tian; Huang & Wang, 2015Junaphaenops Ueno, 1997Junnanotrechus Ueno & Yin, 1993Kettlotrechus Townsend, 2010Kosswigia Jeannel, 1947Kozlovites Jeannel, 1935Kupetrechus Larochelle & Larivi\u00e8re, 2007Kurasawatrechus Yoshida & Nomura, 1952Kusumia Ueno, 1952Lanxangaphaenops Deuve, 2012Laoblemus Ueno, 2006Laosaphaenops Deuve, 2000Lessinodytes Vigna Taglianti, 1982Libotrechus Ueno, 1998Luoxiaotrechus Tian & Yin, 2013Luyatrechus M.Etonti & Mateu, 2000Luzonotrechus Ueno, 1979Mamesdytes Trezzi, 2003Masuzoa Ueno, 1960Masuzonoblemus Ueno, 1989Mayaphaenops Vigna Taglianti, 1977Meganophthalmus Kurnakov, 1959Mexaphaenops Bolivar y Pieltain, 1942Mexitrechus Barr, 1982Miaotrechus Tian; Chen & Ma, 2020Microblemus Ueno, 2007Mimanillus B.Moore, 1972Mimotrechus B.Moore, 1972Minimaphaenops Deuve, 2000Minosaphaenops Queinnec, 2008Minutotrechus Deuve & Kavanaugh, 2016Nannotrechus Winkler, 1926Nanotrechiama Belousov & Kabak, 2018Neaphaenops Jeannel, 1920Nelsonites Valentine, 1952Neotrechus G.M\u00fcller, 1913Nesaeoduvalius Casale & Giachino, 2016Nesiotrechus Ueno, 1995Nipponaphaenops Ueno, 1971Nototrechus B.Moore, 1972Nunbergites Pawlowski & Stachowiak, 1991Omalodera Blanchard, 1842Oodinotrechus Ueno, 1998Oroblemites Ueno & Pawlowski, 1981Oroblemus Ueno & Yoshida, 1966Orotrechus G.M\u00fcller, 1913Oxytrechus Jeannel, 1927Panaphaenops Tian; Huang & Ma, 2021Paracimmerites Belousov, 1998Paraphaenops Jeannel, 1916Paratrechiotes Ueno, 1995Paratrechus Jeannel, 1920Petraphaenops Delic; Kapla & Colla, 2019Pheggomisetes Knirsch, 1923Pilosaphaenops Deuve & Tian, 2008Plesioaphaenops Deuve & Tian, 2011Pogonoschema Jeannel, 1927Pontodytes Casale & Giachino, 1989Porocimmerites Belousov, 1998Protrechiama Belousov & Kabak, 2003Pseudanophthalmus Jeannel, 1920Pseudaphaenops Winkler, 1912Pseudocnides Jeannel, 1927Pseudotrechisibus Mateu & Belles, 1982Puertrechus Belousov & Kabak, 2014Putzeysius Jeannel, 1962Qianaphaenops Ueno, 2000Qianotrechus Ueno, 2000Queinnectrechus Deuve, 1992Rakantrechus Ueno, 1951Ryugadous Habu, 1950Sardaphaenops Cerruti & Henrot, 1956Satotrechus Ueno, 2006Sbordoniella Vigna Taglianti, 1980Scotoplanetes Absolon, 1913Scototrechus Britton, 1962Shenaphaenops Ueno, 1999Shennongotrechus Ueno, 2004Shenoblemus Tian & Fang, 2020Shilinotrechus Ueno, 2003Shiqianaphaenops Tian, 2016Shuangheaphaenops Tian, 2017Shuaphaenops Ueno, 1999Sichuanotrechus Deuve, 2005Sidublemus Tian & Yin, 2013Sinaphaenops Ueno & Wang, 1991Sinocimmerites Deuve, 2007Sinotrechiama Ueno, 2000Sinotroglodytes Deuve, 1996Sloanella Jeannel, 1927Speotrechus Jeannel, 1922Stevensius Jeannel, 1923Stygiotrechus Ueno, 1958Subilsia Espa\u00f1ol, 1967Sumatrechus Deuve, 2005Superbotrechus Deuve & Tian, 2009Suzuka Ueno, 1956Taiwanotrechus Ueno, 1987Taniatrechus Belousov & Dolzhanski, 1994Tasmanorites Jeannel, 1927Tasmanotrechus B.Moore, 1972Taurocimmerites Belousov, 1998Thalassoduvalius Ueno, 1956Tianeotrechus Tian & Tang, 2016Tianzhuaphaenops Zhao & Tian, 2016Tienmutrechus Suenson, 1957Tonkinaphaenops Deuve, 2013Toshiaphaenops Ueno, 1999Trechepaphiama Deuve & Kavanaugh, 2016Trechepaphiopsis Deuve & Kavanaugh, 2016Trechiama Jeannel, 1927Trechiamiotes Deuve, 1998Trechicomimus Mateu & N\u00e8gre, 1972Trechiella Jeannel, 1927Trechimorphus Jeannel, 1927Trechinotus Jeannel, 1962Trechiotes Jeannel, 1954Trechisibus Motschulsky, 1862Trechistus B.Moore, 1972Trechoblemus Ganglbauer, 1891Trechus Clairville, 1806Trichaphaenops Jeannel, 1916Troglocimmerites Ljovuschkin, 1970Tropidotrechus Jeannel, 1927Typhlotrechus G.M\u00fcller, 1913Uenoaphaenops Tian & He, 2020Uenoites Belousov & Kabak, 2016Uenotrechus Deuve & Tian, 1999Ushijimaella Ueno, 1980Velebitaphaenops Casale & Jalzic, 2012Velesaphaenops S.Curcic et al., 2018Vietotrechus Ueno, 1995Wanhuaphaenops Tian & Wang, 2016Wanoblemus Tian & Fang, 2016Wulongoblemus Ueno, 2007Xenotrechus Barr & Krekeler, 1967Xiangxius Tian; Huang & Li, 2021Xuedytes Tian & Huang, 2017Yalongaphaenops Belousov & Kabak, 2021Yamautidius Ueno, 1957Yanzaphaenops Ueno, 2010Yunotrechus Tian & Huang, 2014Zhijinaphaenops Ueno & Ran, 2002Subtribe Trechodina Jeannel, 1926Amblystogenium Enderlein, 1905Canarobius Machado, 1987Cnides Motschulsky, 1862Cothresia Jeannel, 1964Cyphotrechodes Jeannel, 1926Eotrechodes Ueno; Lafer & Sundukov, 1995Himalotrechodes Ueno, 1981Iberotrechodes Faille et al., 2021Pachydesus Motschulsky, 1864Pachytrechodes Jeannel, 1960Paratrechodes Jeannel, 1926Scaurotrechodes Geginat, 2006Spelaeovulcania Machado, 1987Sporades Fauvel, 1882Thalassophilus Wollaston, 1854Trechobembix Jeannel, 1926Trechodes Blackburn, 1901Trechosia Jeannel, 1926Trechosiella Jeannel, 1960\u2020Antarctotrechus Ashworth & Erwin, 2016\u2020Trechinites Heer, 1862\u2020Trechoides Motschulsky, 1856Tribe Zolini Sharp, 1886Subtribe Merizodontina Sloane, 1920Idacarabus Lea, 1910Maungazolus Larochelle & Larivi\u00e8re, 2017Merizodus Solier, 1849Percodermus Sloane, 1920Pseudoopterus Csiki, 1928Pterocyrtus Sloane, 1920Sloaneana Csiki, 1933Synteratus Broun, 1909Thayerella Baehr, 2016Subtribe Zolina Sharp, 1886Oopterus Gu\u00e9rin-M\u00e9neville, 1841Zolus Sharp, 1886Subfamily Xenaroswellianinae Erwin, 2007[edit]Xenaroswelliana Erwin, 2007Subfamily \u2020Conjunctiinae Ponomarenko, 1977[edit]\u2020Conjunctia Ponomarenko, 1977Subfamily \u2020Protorabinae Ponomarenko, 1977[edit]\u2020Aethocarabus Ren; Lu & Guo, 1995\u2020Atrirabus Hong & Wang, 1990\u2020Cordorabus Ponomarenko, 1977\u2020Cretorabus Ponomarenko, 1977\u2020Denudirabus Ren; Lu & Guo, 1995\u2020Lirabus Hong, 1992\u2020Lithorabus Ponomarenko, 1977\u2020Magnirabus Hong & Wang, 1990\u2020Mesorabus Ponomarenko, 1977\u2020Nebrorabus Ponomarenko, 1989\u2020Ovrabites Ponomarenko, 1977\u2020Penecupes Ren, 1995\u2020Protorabus Ponomarenko, 1977Not Assigned to Subfamily\u2020Agatoides Motschulsky, 1856\u2020Amphoxyne Bode, 1953\u2020Carabites Heer, 1852\u2020Cavicarabus Hong, 1991\u2020Conexicoxa Lin, 1986\u2020Cymatopterus Lomnicki, 1894\u2020Fangshania Hong, 1981\u2020Glenopterus Heer, 1847\u2020Hebeicarabus Hong, 1983\u2020Megacarabus Hong, 1983\u2020Meileyingia Hong, 1987\u2020Miocarabus Hong, 1983\u2020Neothanes Scudder, 1890\u2020Procarabus Oppenheim, 1888\u2020Prosynactus Bode, 1953\u2020Shanwangicarabus Hong, 1985\u2020Sinis Heer, 1862\u2020Sinocalosoma Hong & Wang, 1986\u2020Sinocaralosoma Hong, 1984\u2020Sunocarabus Hong, 1987\u2020Tauredon Handlirsch, 1910\u2020Wuchangicarabus Hong, 1991\u2020Xishanocarabus Hong, 1984\u2020Yunnanocarabus Lin, 1977References[edit]"},{"@context":"http:\/\/schema.org\/","@type":"BreadcrumbList","itemListElement":[{"@type":"ListItem","position":1,"item":{"@id":"https:\/\/wiki.edu.vn\/en\/wiki40\/#breadcrumbitem","name":"Enzyklop\u00e4die"}},{"@type":"ListItem","position":2,"item":{"@id":"https:\/\/wiki.edu.vn\/en\/wiki40\/list-of-ground-beetle-genera\/#breadcrumbitem","name":"List of ground beetle genera"}}]}]